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Paul, you have sharp eyes but in the paper they say:

"The deletion of only two genes prevented the production of an active EDF (fig. S11): zwf encoding NNWDN (D = Asp) and ygeO encoding NNWN. The zwf product, carrying the sequence NNWDN, may be the precursor of EDF, and a subsequent amidation step may generate the full NNWNN sequence. Amidation may occur either before or after the cleavage of the precursor by one of E. coli proteases."


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